This species was first described by Devillers-Terschuren and Devillers from Monte Argentaria, Italy
in 2000. Its name means "of the western sunset" which we can only assume refers to the often reddish
hues of the labellum.
Its true status as a species is a matter of some disagreement amongst experts with some believing that
O. hespera is synonymous with O. sulcata. There is little doubt that despite the two species being placed
in different groups they are outwardly very alike. Among the more obvious similarities are :- both being very small, both usually exhibiting the previously mentioned red hues and both having tightly reflexed
lateral lobes that give the overall labellum a cylindrical look.
The range of O. hespera is uncertain but is thought to be mid western Italy centred on the Grosetto region
of Tuscany and Monte Argentaria in particular. In this area its unlikely to be confused with any other
Fusca and in fact the only species with which it can regularly come into contact is the extremely rare
O. lucifera. The clear differences in flower size, shape and colouration are sufficient to differentiate
the two species and in addition, the flowering periods barely overlap with O. hespera being the later
( mid April to May ).
Its a quite variable species in terms of colouration and speculum pattern but the large stigmatic cavity,
and extensive central groove to the lip are generally consistent features. The photograph's are all from
Monte Argentaria and date from the first week of May.
.